A lamella, or gill, is a papery hymenophore rib under the cap of some mushroom species, most often but not always agarics. The gills are used by the mushrooms as a means of spore dispersal, and are important for species identification. The attachment of the gills to the stem is classified based on the shape of the gills when viewed from the side, while color, crowding and the shape of individual gills can also be important features. Additionally, gills can have distinctive microscopic or macroscopic features. For instance, Lactarius species typically seep latex from their gills.
It was originally believed that all gilled fungi were Agaricales, but as fungi were studied in more detail, some gilled species were demonstrated not to be. It is now clear that this is a case of convergent evolution (i.e. gill-like structures evolved separately) rather than being an anatomic feature that evolved only once. The apparent reason that various basidiomycetes have evolved gills is that it is the most effective means of increasing the ratio of surface area to mass, which increases the potential for spore production and dispersal.
Other groups of fungi to bear gills include:
Morphologically, gills are classified according to their attachment to the stem: